This inventory was conducted from 1914 to 1922 shortly after the end of the Little Ice Age and at the leading edge of the severe droughts of the 1920–1930s (Keen, 1937). Current and projected climates are generally drier and warmer than the climate of the centuries preceding this inventory JQ1 purchase and during which the inventoried trees would have established and survived. Longer, drier summers are projected for the Pacific Northwest (Salathé et al., 2010) along with increases in fire frequency (McKenzie et al., 2004). Correlation of sediment records with reconstructed climate show increased biomass burning with

increases in temperature and drought (Marlon et al., 2012). Increases in length of fire season and the size (Westerling et al., 2006) and severity (Miller et al., 2009) of wildfires have already been observed. Fortunately, treatments suggested to increase mean diameter, shift species composition to favor drought- and fire-tolerant species, and restore spatial heterogeneity

Lumacaftor chemical structure in dry forests under current climates are largely consistent with treatments appropriate to at least partially prepare dry forests to deal with expected changes in climate and disturbance regimes (Franklin et al., 1991, Spies et al., 2010a, Spies et al., 2010b, Stephens et al., 2010, Chmura et al., 2011 and Peterson et al., 2011). Historical conditions in the dry forests of south-central Oregon are uniquely documented in an extensive timber inventory (“cruise”) conducted between 1914 and 1922 by the Bureau of Indian Affairs (BIA) on the former Klamath Indian Reservation (now a part of the Fremont-Winema National Forskolin manufacturer Forest). The forested area of the reservation was sampled at 10–20% intensity using a systematic

grid consisting of one or two 1.6 ha transects per quarter-quarter section (16.2 ha). Transect location was tied to documented survey points of the Bureau of Land Management Public Land Survey System (BLM PLSS). Live conifers at least 15 cm dbh were tallied by species and diameter class. This archived inventory represents a large and systematic sample of historical forest composition and structure over hundreds of thousands of hectares, which complements existing historical records and reconstructions for this area (Table 6). Similar inventory records from other forested areas have been used to understand historical conditions and to validate reconstructions of reference conditions in the central Sierra Nevada in California (Scholl and Taylor, 2010 and Collins et al., 2011) and in Australia (Whipp et al., 2010). Our focus in this paper is on the historical range of variability in structure and composition of dry forests growing on ponderosa pine and mixed-conifer habitat types (potential vegetation types) in three large segments (117,672 ha total) of the former Klamath Indian Reservation as recorded in the 1914–1922 timber inventory. In addition to documenting the historical structure and composition at the stand (1.